Been too long since I've done a parasite post, so...
Lungworm! And Fungus Cannon!
Dictyocaulus is a genus of nematode worms, parasitic upon large herbivores including cattle, elk, deer, horses, and donkeys. They spend most of their life in the lungs, causing the disease 'Husk'. Their eggs, however, get coughed up and swallowed, and hatch into L1
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While working on this research, we began playing with nematodes as stand-ins for the lungworms. Like the lungworms, they crawl all over the place, including all over the Pilobolus stalks.
It is not clear if lungworms actually end up hitching a ride on the spores or not. And if they do, it's not clear if they purposefully climb up the stalks or just end up there on accident. Hopefully further research will yield some more answers--and some more WTF moments ;)
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Since the spores shoot towards the sun, they have to be pointing towards the sun, which will at least get the critters off the cowpatty itself. There might be some weather-related assistance in there somewhere, too, if a breeze is blowing at launch time.
I'm also guessing the 55 MPH launch was developed specifically to overcome viscous vs inertia.
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